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Medway, Lord 1965. Niah Cave animal bone, VIII: Rhinoceros in late quaternary Borneo. Sarawak Museum Journal 12 (25/26): 77-82, pl. 21.

Niah Cave animal bone, VIII: Rhinoceros in late quaternary Borneo

Note
Location Borneo Subject Distribution Species Sumatran Rhino (sumatrensis)

Specimen unspecified. Locality: Sandakan. Collected by: W.B. Pryer, 1886. In Natural History Museum, London, United Kingdom.

Note
Location Borneo Subject Distribution Species Sumatran Rhino (sumatrensis) Year 1886

Specimen unspecified. Locality: Sandakan. Collected by: W.B. Pryer, 1886. In Natural History Museum, London, United Kingdom.

Note
Location Borneo Subject Distribution Species Sumatran Rhino (sumatrensis)

Mounted head. Locality: Sabah, S. Tingkayu. In coll. National Museum, Singapore

Note
Location Borneo Subject Distribution Species Sumatran Rhino (sumatrensis) Year 1886

Specimen unspecified. Locality: Sandakan. Collected by: W.B. Pryer, 1886. In Natural History Museum, London, United Kingdom.

Note
Location Borneo Subject Distribution Species Sumatran Rhino (sumatrensis)

Mounted head. Locality: Sabah, S. Tingkayu. In coll. National Museum, Singapore

Note
Location Borneo Subject Distribution Species Sumatran Rhino (sumatrensis) Year 1895

The second previous collection of rhinoceros remains from Sarawak consists of a series of bones together with two upper molars, recovered from gold workings in an alluvial cave deposit in the upper Sarawak river, presumably at or near Ban. These were presented to the British Museum (Natural History) by Dr. P. Lutley Sclater. The two teeth were provisionally referred to Rh. sondaicus by Lydekker (1886, p.129), but were entered in the British Museum register for 1895 (reg. no. M1986) as sumatrensis (A. J. Sutcliffe, in litt.). Hooijer (1946) his pointed out that the associated post-cranial bones have consistently been referred to sumatrensis.

Note
Location Borneo Subject Distribution Species Sumatran Rhino (sumatrensis)

The second previous collection of rhinoceros remains from Sarawak consists of a series of bones together with two upper molars, recovered from gold workings in an alluvial cave deposit in the upper Sarawak river, presumably at or near Ban. These were presented to the British Museum (Natural History) by Dr. P. Lutley Sclater. The two teeth ivere provisionally referred to Rh. sondaicus by Lydekker (1886, p.129), but were entered in the British Museum register for 1895 (reg. no. M1986) as sumatrensis (A. J. Sutcliffe, in litt.). Hooijer (1946) his pointed out that the associated post-cranial bones have consistently been referred to sumatrensis. During the preparation of his paper on prehistoric and fossil rhinoceroses from the Sunda region. Dr. Hooijer was unable to examine these controversial molars (Hooijer, 1946, p.10). However. a photograph has now been made available through the kindness of Dr. A. J. Sutclifre, Department of Palaeontology and is published here (see Plate XXI). The teeth are seen to be first upper molars, one from the right and one from the left jaw, exhibiting different degrees of wear and presumably from different animals. The dimensions, measured from the photograph, are small compared with the figures for recent D. sumatrensis given by Hooijer (1946) and would be exceptional for Rh. sondaicus. In addition, certain morphological features peculiar to sumatrensis are clearly visible in the photographs. Firstly, in the more worn tooth in particular a distinct bend in the enamel at the antero-lingual margin of the protoloph is seen, indicating the presence of the ?protocone fold' of Hooijer (1946. p.11). On the less worn tooth the protocone fold is also present, although less clearly shown in the photograph. on the other hand, the relatively unworn condition of the latter tooth permits a second diagnostic character to be seen viz., in the words of Hooijer (1946), that the crochet ?springs off from the metaloph below the upper margin', rather than originating at the apex of the metaloph as in sondalcus. Together, these characters confirm the identification of the molars as D. sumatrensis.

Note
Location Borneo Subject Distribution Species Sumatran Rhino (sumatrensis)

Borneo, Sarawak - teeth mentioned by Busk In addition to the material from Niah, fossil or subfossil rhinoceros remains have previously been recorded also from south-western Sarawak. The first to come to light were two teeth sent to Sir Charles Lyell by Rajah James Brooke, and discussed by G. Busk (1869). These teeth were identified by Busk as right and left second upper molars, evidently belonging to the same individual, both consisting only of parts of uncrupted crowns in what the author refers to as the ?germ' state. Busk considered that they could be attributed to a very young Rh. sondaicus. However, Hooijer subsequently re-examined Busk's evidence and concluded, on the basis of his figures, that the molars were in fact those of D. sumatrensis (Hooijer, 1945).

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