Payne, J.; Andau, M. 1991. Large mammals in Sabah. In Kiew, R. (ed), The state of nature conservation in Malaysia (Kuala Lumpur, Malayan Nature Society). pp. 177-183.
Fraser, D.; Gilbert, M.; Rybczynski, N.; Dawson, M.R. 2025. Post-Eocene rhinocerotid dispersal via the North Atlantic. Vertebrate Anatomy, Morphology, Palaeontology 13 (Canadian Society of Vertebrate Palaeontology 2025 Abstracts): 21.