Williams, B.; Campbell, J.; Kendall, C.; Tuttle, J.; Lynch, E.C. 2023. Exploring the behaviors and social preferences of a large, multigenerational herd of zoo-housed Southern white rhinoceros (Ceratotherium simum simum). Zoo Biology 42 (4): 476-489. doi.org/10.1002/zoo.21758
Sinnett, S.; Tuttle, J.; Hinson, A.; Trull, T.; Weatherly, S.; Batot, L.; North Carolina Zoo, Asheboro, North Carolina 2022. Preparing 0.1 Southern White rhino for a dental procedure. Proceedings of the 47th Annual AAZK National Conference Toronto, Canada Oct. 13-17: poster.
Tuttle, J. 2017. Dental surgery for geriatric southern white rhino. Proceedings of the International Rhino Keeper (IRKA) Workshop held in Denver, August 2017.
Tuttle, J. 2015. Which reward (North Carolina Zoo). The Crash April 2015: 2-3.
Tuttle, J. 2014. Rhino open mouth behavior. The Crash January 2014: 6.
Tuttle, J. 2007. Facility focus – the Caldwell Zoo. The Crash 2 (2): 2-3, figs. 1-9.






