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Hammond, E.E.; Penfold, L.M.; Metrione, L.C. 2026 . Preliminary investigation: Down-regulation of testosterone by an anti-GnRH immunotherapeutic (Improvest®) in a Southern White Rhinoceros (Ceratotherium simum simum) bull. Theriogenology 8 (100156): 1-4. doi.org/10.1016/j.therwi.2026.100156
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Stoops, M.; Moresco, A.; Penfold, L.M.; Gillis, J.D.; Wojtusik, J.; Metrione, L.C.; O’Brien, J.K. 2021 . Overcoming reproductive disorders in female greater one-horned rhinoceros (Rhinoceros unicornis) to improve artificial insemination and natural breeding success. Clinical Theriogenology 13 (4) Dec: 394-401. doi.org/10.58292/ct.v13.9317
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Penfold, L.M. 2011 . An overview of the assisted reproduction and genome banking activities of White Oak Conservation Center, Yulee, FL, in the service of species conservation. International Zoo Yearbook 45 (1): 154-159.
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Christensen, B.W.; Troedsson, M.; Young, L.J.; Oliva, M.; Penfold, L.M. 2009 . Effects of sociosexual environment on serum testosterone in captive male African rhinoceros. Theriogenology 71: 1105-1111, tables 1-2.
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Metrione, L.C.; Penfold, L.M.; Waring, G.H. 2007 . Social and spatial relationships in captive southern white rhinoceros (Ceratotherium simum simum). Zoo Biology 26 (4): 487-502, figs. 1-3, tables 1-2.
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Oliva, M.; Steele, V.; President, T.; Penfold, L.M. 2007 . Longitudinal serum progesterone analysis to assess reproductive success in female White rhinoceros (Ceratotherium simum). Proceedings of the AAZV, AAWV, AZA/NAG Joint Conference 2007: 240.
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Christensen, B.W.; Troedsson, M.; Little, R.; Oliva, M.; Penfold, L.M. 2006 . Invasive and non-invasive methods of hormone monitoring in captive wildlife. In: Rhino Keeper Workshop 2006 . Proceedings of the Fourth Rhino Keepers workshop 2005 at Columbus, Ohio. Columbus, Rhino Keepers Workshop. p. 1.
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Christensen, B.W.; Troedsson, M.; Young, L.J.; Penfold, L.M. 2006 . Exogenous influences on serum testosterone concentrations in male African rhinoceros species. Proceedings Phi Zeta Research Emphasis Day: p. 1.
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Myers, D.; Penfold, L.M. 2004 . Early pregnancy detection in rhinoceros species using serum glycoproteins. Proceedings of the American Association of Zoo Veterinarians (AAZV) 2004: 604-605.
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Christensen, B.W.; Troedsson, M.; Oliva, M.; MacPherson, M.; Pozor, M.; Penfold, L.M. 2004 . Exogenous influences on serum testosterone concentration in captive black rhinoceros (Diceros bicornis). Proceedings of the American Association of Zoo Veterinarians (AAZV) 2004: 536.
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