Harley, E.H.; De Waal, M.; Murray, Shane; Ryan, C. O’ 2016. Comparison of whole mitochondrial genome sequences of northern and southern white rhinoceroses (Ceratotherium simum): the conservation consequences of species definitions. Conservation Genetics 17: 1285–1291.
Robinson, T.J.; Trifonov, V.; Espie, I.; Harley, E.H. 2005. Interspecific hybridisation in rhinoceroses: confirmation of a black x white rhinoceros hybrid by karyotype, fluorescence in sity hybridisation (FISH) and microsatellite analysis. Conservation Genetics 6: 141-145, figs. 1-2, table 1.
Harley, E.H.; Baumgarten, I.; Cunningham, J.; Ryan, C. O’ 2005. Genetic variation and population structure in remnant populations of black rhinoceros, Diceros bicornis, in Africa. Molecular Ecology 14: 2981-2990, fig. 1, tabs. 1-6.
Weber, B.; Paglia, D.E.; Harley, E.H. 2004. Elevated free tyrosine in rhinoceros erythrocytes. Comparative Biochemistry and Physiology 138A: 105-109, figs. 1-4.
Harley, E.H.; Matshikiza, M.; Robson, P.; Weber, B. 2004. Red blood cell metabolism show major anomalies in Rhinocerotidae and Equidae, suggesting a novel role in general antioxidant metabolism. International Congress Series 1275: 334-340, figs. 1-4, tables 1-2.
Robinson, T.J.; Trifonov, V.; F. Yang, F.; Espie, I.; Harley, E.H. 2003. Correspondence of chromosome segments in low numbered (Equidae) and high numbered (Rhinocerotidae) Perissodactyla, and genetic confirmation of an F1 black x white rhinoceros by FISH and microsatellite analysis. Annales de Genetique 46 (2/3): 290.
Weber, B.; Paglia, D.E.; Harley, E.H. 2002. Red cell metabolism in the black rhinoceros: relevance to haemolytic disease. p. 181.
Ryan, C. O’; Cunningham, J.; Harley, E.H. 2002. Microsatellite analysis of African black rhinoceros (Diceros bicornis) to determine genetic diversity and population structure.
Paglia, D.E.; Weber, B.; Baumgarten, I.; Harley, E.H. 2001. Radiometric assessment of hexose monophosphate shunt capacity in erythrocytes of rhinoceroses. American Journal of Veterinary Research 62 (7): 1113-1117, fig. 1, tables 1-2.
Weber, B.; Paglia, D.E.; Harley, E.H. 2001. Red cell metabolism in the black rhinoceros: relevance to haemolytic disease. p. 41.
O’Ryan, C.; Cunningham, J.; Baumgarten, I.; Harley, E.H. 2001. Microsatellite analysis of African black rhinoceros (Diceros bicornis) to determine genetic diversity and population structure.
Cunningham, J.; Harley, E.H.; O’Ryan, C. 1999. Isolation and characterization of microsatellite loci in black rhinoceros (Diceros bicornis). Electrophoresis 20 (8): 1778-1780, tables 1-2.
Harley, E.H.; Paglia, D.E.; Weber, B. 1997. Oxidative damage and purine metabolism: investigation of haemolytic anaemia in the black rhinoceros. Clinical Biochemistry 30 (4): 259.
Ryan, C. O’; Flamand, J.R.B.; Harley, E.H. 1994. Mitochondrial DNA variation in black rhinoceros (Diceros bicornis): conservation management implications. Conservation Biology 8 (2): 495-500, figs. 1-2, tables 1-3.
Harley, E.H.; Ryan, C. O’ 1994. Use of molecular genetics in rhinoceros conservation.
Ryan, C. O’; Harley, E.H. 1993. Comparison of mitochondrial DNA in black and white rhinoceroses. Journal of Mammalogy 74 (2): 343-346, fig. 1, table 1.
Harley, E.H.; Ryan, C. O’ 1993. Molecular genetic studies of southern African rhinoceros.






