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References by this author
Handa, N.; Nishioka, Y.; Kusaka, S.; Duangkrayom, J.; Naksri, W.; Jintasakul, P.; Kunimatsu, Y. 2026. The Late Miocene Rhinocerotidae from the Phra Phut sandpit in northeastern Thailand: implication for biostratigraphy and palaeobiogeography. Palaeobiodiversity and Palaeoenvironments. doi.org/10.1007/s12549-026-00714-4
Handa, N.; Taru, H. 2025. Taxonomic revision of a late Miocene rhinoceros from Japan with an overview of Brachypotherium from East Asia. Historical Biology 37 (8): 1874-1879. doi.org/10.1080/08912963.2025.2456950
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Longuet, M.; Handa, N.; Zin Maung Maung Thein; Thaung Htike; Man-Thit-Nyein; Takai, Masanaru 2024. Post-cranial remains of Rhinocerotidae from the Neogene of central Myanmar: morphological descriptions and comparisons with ratios. Historical Biology 37 (8): 1859-1873. doi.org/10.1080/08912963.2024.2408617
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Handa, N.; Takahashi, K. 2024. A review of the taxonomy, biostratigraphy and paleobiogeography of Plio-Pleistocene rhinoceroses in Japan. Journal of Asian Earth Sciences 276 (106357): 1-15. doi.org/10.1016/j.jseaes.2024.106357
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Handa, N.; Izuho, M.; Takahashi, K.; Iizuka, F.; Tsogtbaatar, B.; Gunchinsuren, B.; Odosuren, D.; Ishitseren, L. 2022. The woolly rhinoceros (Coelodonta antiquitatis) from Ondorkhaan, eastern Mongolia. Boreas 51 (3): 584-605, 10 figs, 1 tab. doi.org/10.1111/bor.12582
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Handa, N.; Tsubamoto, T. 2021. Reappraisal of a supposed chalicotheriid perissodactyl femur from the Pliocene of Yenangyoung, central Myanmar. Journal of Fossil Research 54 (1): 11-14.
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Handa, N.; Nishioka, Y.; Duangkrayom, J.; Jintasakul, P. 2021. Brachypotherium perimense (Perissodactyla, Rhinocerotidae) from the Miocene of Nakhon Ratchasima, Northeastern Thailand, with comments on fossil records of Brachypotherium. Historical Biology 33 (9): 1642-1660. doi.org/10.1080/08912963.2020.1723578
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Handa, N.; Kato, Takafumi 2020. A Pliocene rhinocerotid (Mammalia, Perissodactyla) from Ajimu, Oita Prefecture, southwestern Japan, with comments on the Japanese Pliocene rhinocerotid fossil records. Paläontologische Zeitschrift 94: 759-768.
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Handa, N. 2019. Reassessment of a Pleistocene rhinocerotid (Mammalia, Perissodactyla) from Aira, Kagoshima, Southwestern Japan. Paleontological Research 23 (1): 55-64, 5 figs, 1 tab. doi.org/10.2517/2018PR009
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Handa, N.; Nakatsukasa, M.; Kunimatsu, Y.; Nakaya, H. 2019. Additional specimens of Diceros (Perissodactyla, Rhinocerotidae) from the Upper Miocene Nakali Formation in Nakali, central Kenya. Historical Biology 31 (2): 262-273. doi.org/10.1080/08912963.2017.1362560
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Handa, N. 2019. A review of the miocene rhinoceroses from japan, and paleobiogeographic implications. Abstracts Society of Vertebrate Paleontology 79th Annual Meeting: 113-114.
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Handa, N.; Kohno, N.; Kudo, Y. 2019. Reappraisal of a middle Pleistocene rhinocerotid (Mammalia, Perissodactyla) from the Matsugae Cave, Fukuoka Prefecture, southwestern Japan. Historical Biology 33 (2): 218-229, 5 figs, 3 tabs. doi.org/10.1080/08912963.2019.1604699
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Handa, N.; Nakatsukasa, M.; Kunimatsu, Y.; Nakaya, H. 2018. Brachypotherium (Perissodactyla, Rhinocerotidae) from the late Miocene of Samburu Hills, Kenya. Geobios 51 (5): 391-399.
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Handa, N.; Nakatsukasa, M.; Kunimatsu, Y.; Nakaya, H. 2017. Additional specimens of Diceros (Perissodactyla, Rhinocerotidae) from the Upper Miocene Nakali Formation in Nakali, central Kenya. Historical Biology 31 (2): 262-273, 6 figs, 3 tabs. doi.org/10.1080/08912963.2017.1362560
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Handa, N.; Nakatsukasa, M.; Kunimatsu, Y.; Nakaya, H. 2017. A new Elasmotheriini (Perissodactyla, Rhinocerotidae) from the upper Miocene of Samburu Hills and Nakali, northern Kenya. Geobios 50 (3): 197-209.
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Handa, N.; Takechi, Y. 2017. A Pleistocene rhinocerotid (Mammalia, Perissodactyla) from the Bisan-Seto area, western Japan. Journal of the Geological Society of Japan 123 (6): 433-441, 4 figs, 2 tabs. doi.org/10.5575/geoscoc.2017.0004
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Handa, N.; Izuho, M.; Takahashi, K.; Iizuka, F.; Tsogtbaatar, B.; Gunchinsuren, B.; Odosuren, D.; Ishitseren, L. 2017. Discovery of a Late Pleistocene rhinocerotid fossil from Ondorkhaan, eastern Mongolia. Journal of the Geological Society of Japan 123 (12): v-vi. doi.org/10.5575/geosoc.2017.0058
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Handa, N. 2016. Fossil rhinoceroses from the Late Miocene of Kenya: Miocene faunal changes of the Rhinocerotidae (Mammalia, Perissodactyla) in sub-Saharan East Africa. Report to Kagoshima University. pp. 1-28.
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Handa, N.; Pandolfi, L. 2016. Reassessment of the Middle Pleistocene Japanese Rhinoceroses (Mammalia, Rhinocerotidae) and paleobiogeographic implications. Paleontological Research, The Palaeontological Society of Japan 20 (3): 247-260, 4 figs, 3 tabs. doi.org/10.2517/2015PR034
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Handa, N. 2015. A Pleistocene rhinocerotid (Mammalia, Perissodactyla) from Yage, Shizuoka Prefecture, central Japan. Paleontological Research, The Palaeontological Society of Japan; 19 (2): 139-142, 3 figs, 1 tab [http://www.bioone.org/doi/full/10.2517/2014PR033].
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Handa, N.; Nakatsukasa, M.; Kunimatsu, Y.; Nakaya, H.; Tsubamoto, T. 2015. New specimens of Chilotheridium (sic) (Perissodactyla, Rhinocerotidae) from the Upper Miocene Namurungule and Nakali Formations, Northern Kenya. Paleontological Research, The Palaeontological Society of Japan 19 (3): 181-194, 5 figs. http://www.bioone.org/doi/full/10.2517/2014PR035
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Nakaya, H.; Nakatsukasa, M.; Kunimatsu, Y.; Handa, N. 2012. The Late Miocene Elasmotheriini (Rhinocerotidae, Mammalia) from the Namurungule and Nakali Formations of northern Kenya. Japan Geoscience Union Meeting 2012, Chiba, Japan: 1.
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Miyata, Kazunori; Saegusa, H.; Nakaya, H.; Ugai, H.; Kato, Takafumi; Handa, N.; et al. 2012. Early Miocene rhinocerotid specimen from the Takashima Formation, Takashima island (Taka Island), Matsuura City, Nagasaki Prefecture.. Abstracts of the 161st Regular Meeting of the Paleontological Society of Japan, Gunma, Japan 2012: 55.
Handa, N.; Nakaya, H.; Nakatsukasa, M.; Kunimatsu, Y. 2012. New specimens of Elasmotheriini (Rhinocerotidae, Perissodactyla) from the Namurungule and Nakali Formations (Early Late Miocene) of Northern Kenya. Journal of Vertebrate Paleontology - 72nd Annual Meeting of the Society of Vertebrate Paleontology, Raleigh, NC, USA, October 17-20, 2012 (abstract); 32 (Sup. 2): 206.
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