Longuet, M.; Handa, N.; Zin-Maung-Maung-Thein; Thaung-Htike; Man-Thit-Nyein; Takai, Masanaru 2024. Post-cranial remains of Rhinocerotidae from the Neogene of central Myanmar: morphological descriptions and comparisons with ratios. Historical Biology 37 (8): 1-10. doi.org/10.1080/08912963.2024.2408617
Handa, N.; Izuho, M.; Takahashi, K.; Iizuka, F.; Tsogtbaatar, B.; Gunchinsuren, B.; Odosuren, D.; Ishitseren, L. 2022. The woolly rhinoceros (Coelodonta antiquitatis) from Ondorkhaan, eastern Mongolia. Boreas 51 (3): 584-605, 10 figs, 1 tab. doi.org/10.1111/bor.12582
Handa, N.; Tsubamoto, T. 2021. Reappraisal of a supposed chalicotheriid perissodactyl femur from the Pliocene of Yenangyoung, central Myanmar. Journal of Fossil Research 54 (1): 11-14.
Handa, N.; Kato, Takafumi 2020. A Pliocene rhinocerotid (Mammalia, Perissodactyla) from Ajimu, Oita Prefecture, southwestern Japan, with comments on the Japanese Pliocene rhinocerotid fossil records. Paläontologische Zeitschrift 94: 759-768.
Handa, N. 2019. Reassessment of a Pleistocene rhinocerotid (Mammalia, Perissodactyla) from Aira, Kagoshima, Southwestern Japan. Paleontological Research 23 (1): 55-64, 5 figs, 1 tab - [. doi.org/10.2517/2018PR009]
Handa, N.; Nakatsukasa, M.; Kunimatsu, Y.; Nakaya, H. 2019. Additional specimens of Diceros (Perissodactyla, Rhinocerotidae) from the Upper Miocene Nakali Formation in Nakali, central Kenya. Historical Biology 31 (2): 262-273. doi.org/10.1080/08912963.2017.1362560
Handa, N. 2019. A review of the miocene rhinoceroses from japan, and paleobiogeographic implications. Abstracts Society of Vertebrate Paleontology 79th Annual Meeting: 113-114.
Handa, N.; Kohno, N.; Kudo Y. 2019. Reappraisal of a middle Pleistocene rhinocerotid (Mammalia, Perissodactyla) from the Matsugae Cave, Fukuoka Prefecture, southwestern Japan. Historical Biology 33 (2): 218-229, 5 figs, 3 tabs. doi.org/10.1080/08912963.2019.1604699
Handa, N.; Nakatsukasa, M.; Kunimatsu, Y.; Nakaya, H. 2018. Brachypotherium (Perissodactyla, Rhinocerotidae) from the late Miocene of Samburu Hills, Kenya. Geobios 51 (5): 391-399.
Handa, N.; Nakatsukasa, M.; Kunimatsu, Y.; Nakaya, H. 2017. Additional specimens of Diceros (Perissodactyla, Rhinocerotidae) from the Upper Miocene Nakali Formation in Nakali, central Kenya. Historical Biology 31 (2): 262-273, 6 figs, 3 tabs [. doi.org/10.1080/08912963.2017.1362560]
Handa, N.; Nakatsukasa, M.; Kunimatsu, Y.; Nakaya, H. 2017. A new Elasmotheriini (Perissodactyla, Rhinocerotidae) from the upper Miocene of Samburu Hills and Nakali, northern Kenya. Geobios 50 (3): 197-209.
Handa, N.; Takechi, Y. 2017. A Pleistocene rhinocerotid (Mammalia, Perissodactyla) from the Bisan-Seto area, western Japan. Journal of the Geological Society of Japan 123 (6): 433-441, 4 figs, 2 tabs [. doi.org/10.5575/geoscoc.2017.0004]
Handa, N.; Izuho, M.; Takahashi, K.; Iizuka, F.; Tsogtbaatar, B.; Gunchinsuren, B.; Odosuren, D.; Ishitseren, L. 2017. The woolly rhinoceros (Coelodonta antiquitatis) from Ondorkhaan, eastern Mongolia [in Japanese]. Journal of the Geological Society of Japan 123(12): V-VI.
Handa, N.; Izuho, M.; Takahashi, K.; Iizuka, F.; Tsogtbaatar, B.; Gunchinsuren, B.; Odosuren, D.; Ishitseren, L. 2017. Discovery of a Late Pleistocene rhinocerotid fossil from Ondorkhaan, eastern Mongolia. Journal of the Geological Society of Japan 123 (12): v-vi. doi.org/10.5575/geosoc.2017.0058
Handa, N. 2016. Fossil rhinoceroses from the Late Miocene of Kenya: Miocene faunal changes of the Rhinocerotidae (Mammalia, Perissodactyla) in sub-Saharan East Africa. Report to Kagoshima University. pp. 1-28.
Handa, N.; Pandolfi, L. 2016. Reassessment of the Middle Pleistocene Japanese Rhinoceroses (Mammalia, Rhinocerotidae) and paleobiogeographic implications. Paleontological Research, The Palaeontological Society of Japan; 20 (3): 247-260, 4 figs, 3 tabs [doi: http://dx.doi.org/10.2517/2015PR034].
Handa, N. 2015. A Pleistocene rhinocerotid (Mammalia, Perissodactyla) from Yage, Shizuoka Prefecture, central Japan. Paleontological Research, The Palaeontological Society of Japan; 19 (2): 139-142, 3 figs, 1 tab [http://www.bioone.org/doi/full/10.2517/2014PR033].
Handa, N.; Nakatsukasa, M.; Kunimatsu, Y.; Tsubamoto, Takehisa; Nakaya, H. 2015. New specimens of Chilotheridium (sic) (Perissodactyla, Rhinocerotidae) from the Upper Miocene Namurungule and Nakali Formations, Northern Kenya. Paleontological Research, The Palaeontological Society of Japan; 19 (3): 181-194, 5 figs [http://www.bioone.org/doi/full/10.2517/2014PR035].
Handa, N.; Nakaya, H.; Nakatsukasa, M.; Kunimatsu, Y. 2012. New specimens of Elasmotheriini (Rhinocerotidae, Perissodactyla) from the Namurungule and Nakali Formations (Early Late Miocene) of Northern Kenya. Journal of Vertebrate Paleontology - 72nd Annual Meeting of the Society of Vertebrate Paleontology, Raleigh, NC, USA, October 17-20, 2012 (abstract); 32 (Sup. 2): 206.
Nakaya, H.; Kato, Takafumi; Handa, N. 2012. Early Miocene rhinocerotid specimen from the "Takashima Formation", Takashima island (Taka Island), Matsuura City, Nagasaki Prefecture..






