Sauspeter, G.; Clauss, M.; Ortmann, S.; Abraham, A.J.; Biddle, R.; Versteege, L.; Przybylo, M. 2025. Feeding management of African rhinos (Ceratotherium simum, Diceros bicornis) in European zoos. Zoo Biology 2025: 1-15. doi.org/10.1002/zoo.70031
Wittwer, A.; Roller, M.; Mueller, D.W.H.; Bertelsen, M.F.; Lackey, L.B.; Steck, B.; Biddle, R.; Versteege, L.; Clauss, M. 2023. Historical development of the survivorship of zoo rhinoceroses – a comparative historical analysis. Zoo Biology 2023: 1-14. doi.org/10.1002/zoo.21793
Radeke-Auer, K.; Wittwer, A.; Aust, J.; Roller, M.; Mueller, D.; Houwald, F. von; Steck, B.; Biddle, R.; Versteege, L.; Clauss, M. 2022. Reproductive non-seasonality in rhinoceroses: A review of the in-situ literature and birth records of ex-situ institutions. Journal of Zoo and Aquarium Research 10 (1): 23-30. doi.org/10.19227/jzar.v10i1.628
Pilgrim, M.; Biddle, R. 2020. EAZA Best Practice Guidelines: Black rhinoceros (Diceros bicornis), edition 2. Amsterdam, EAZA. pp. 1-113.
Pilgrim, M.; Biddle, R. 2015. Eastern black rhinos and Chester Zoo. Paper presented at the 2015 Rhino Keeper Workshop held in Chester, UK: pp. 1-61.
Biddle, R.; Pilgrim, M. 2015. Eastern black rhino EEP (Diceros bicornis michaeli): regional studbook report 2015. Chester Zoo: pp. 1-20.
Biddle, R.; Pilgrim, M. 2014. Eastern black rhino EEP (Diceros bicornis michaeli): regional studbook report 2014. Chester Zoo: pp. 1-27.
Pilgrim, M.; Biddle, R. 2014. EAZA Best Practice Guidelines: Black rhinoceros (Diceros bicornis). Amsterdam, EAZA. pp. 1-85.
Biddle, R.; Pilgrim, M. 2013. Eastern black rhino EEP (Diceros bicornis michaeli): regional studbook report 2013. Chester Zoo: pp. 1-25.
Biddle, R.; Pilgrim, M. 2012. Eastern black rhino EEP (Diceros bicornis michaeli): regional studbook report 2012. Chester Zoo: pp. 1-22.






